Temporal range: 0.28–0 Ma Middle Pleistocene–Present
|Male and female
Homo sapiens sapiens
Homo sapiens (Latin: "wise man") is the binomial nomenclature (also known as the scientific name) for the only extant human species. Homo is the human genus, which also includes Neanderthals and many other extinct species of hominin; H. sapiens is the only surviving species of the genus Homo. Modern humans are the subspecies Homo sapiens sapiens, which differentiates them from what has been argued to be their direct ancestor, Homo sapiens idaltu.
The ingenuity and adaptability of Homo sapiens has led to it becoming the most influential species on Earth; it is currently deemed of least concern on the Red List of endangered species by the International Union for Conservation of Nature.
H. sapiens includes Homo sapiens idaltu, an archaic subspecies of H. sapiens. Extant human populations have historically been divided into subspecies, but since c. the 1980s all extant groups tend to be subsumed into a single subspecies, H. sapiens sapiens.
Some sources show Neanderthals (Homo neanderthalensis) as a subspecies (Homo sapiens neanderthalensis). Similarly, the discovered specimens of the Homo rhodesiensis species have been classified by some as a subspecies (Homo sapiens rhodesiensis), although it remains more common to treat these last two as separate species within the Homo genus rather than as subspecies within H. sapiens.
Since 2010, genetic research has led to the emergence of an intermediate position, characterised by mostly recent African origin plus limited admixture with archaic humans.
The recent African origin of modern humans is the mainstream model that describes the origin and early dispersal of anatomically modern humans. The theory is called the (Recent) Out-of-Africa model in the popular press, and academically the recent single-origin hypothesis (RSOH), Replacement Hypothesis, and Recent African Origin (RAO) model. The hypothesis that humans have a single origin (monogenesis) was published in Charles Darwin's Descent of Man (1871). The concept was speculative until the 1980s, when it was corroborated by a study of present-day mitochondrial DNA, combined with evidence based on physical anthropology of archaic specimens. According to genetic and fossil evidence, archaic Homo sapiens evolved to anatomically modern humans solely in Africa, between 200,000 and 100,000 years ago, with members of one branch leaving Africa by 60,000 years ago and over time replacing earlier human populations such as Neanderthals and Homo erectus. More recently, in 2017, fossils found in Jebel Irhoud (Morocco) suggest that Homo sapiens may have evolved as early as 315,000 years ago, and other evidence suggests that Homo sapiens may have migrated from Africa as early as 270,000 years ago.
The recent single origin of modern humans in East Africa is the near-consensus position held within the scientific community. However, recent sequencing of the full Neanderthal genome suggests Neanderthals and some modern humans share some ancient genetic lineages. The authors of the study suggest that their findings are consistent with Neanderthal admixture of up to 4% in some populations. But the study also suggests that there may be other reasons why humans and Neanderthals share ancient genetic lineages. In August 2012, a study by scientists at the University of Cambridge questioned this conclusion, hypothesising instead that the DNA overlap is a remnant of a common ancestor of both Neanderthals and modern humans. That study however does not explain why only a fraction of modern humans have Neanderthal DNA.
The multiregional origin model, proposed by Milford H. Wolpoff in 1988 provides another explanation for the pattern of human evolution. Multiregional origin holds that the evolution of humanity from the beginning of the Pleistocene 2.5 million years BP to the present day has been within a single, continuous human species, evolving worldwide to modern Homo sapiens sapiens.
The time frame for the evolution of the genus Homo out of the chimpanzee–human last common ancestor is roughly 10 to 2 million years ago, and that of H. sapiens out of Homo erectus roughly 1.8 to 0.2 million years ago.
Scientific study of human evolution is concerned, primarily, with the development of the genus Homo (extant and extinct human species), but usually involves studying other hominids as well, i.e. other "great apes"; these include Australopithecus, an important ancestor of humans, and our current as well as extinct relatives among the Homininae subfamily: chimpanzees, bonobos, gorillas, and the related extinct hominins.
Homo sapiens idaltu, the other known subspecies, is now extinct. Homo neanderthalensis, which became extinct 30,000 years ago, has sometimes been classified as a subspecies, "Homo sapiens neanderthalensis"; genetic studies now suggest that the functional DNA of modern humans and Neanderthals diverged 500,000 years ago.
Similarly, the discovered specimens of the Homo rhodesiensis species have been classified by some as a subspecies, but this classification is not widely accepted.
Anatomically modern humans first appear in the fossil record in Africa about 195,000 years ago (see Omo remains), and studies of molecular biology give evidence that the approximate time of divergence from the common ancestor of all modern human populations was 200,000 years ago. The broad study of African genetic diversity found the ǂKhomani San people to express the greatest genetic diversity among the 113 distinct populations sampled, making them one of 14 "ancestral population clusters". The research also located the origin of modern human migration in southwestern Africa, near the coastal border of Namibia and Angola.
Currently available genetic and archaeological evidence is generally interpreted as supportive of a recent single origin of modern humans in East Africa. However, this is where the near consensus on human settlement history ends, and considerable uncertainty clouds any more detailed aspect of human colonization history.
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